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Whittinghill, M. and Hinton, Claude W. (1950) The Nature of Linkage Variation with Age in Inversion Heterozygotes of Drosophila melanogaster. Proceedings of the National Academy of Sciences of the United States of America, 36 (10). pp. 546-551. ISSN 0027-8424. http://resolver.caltech.edu/CaltechAUTHORS:WHIpnas50